Ayestaran has not yet officially left the club but has had a major fall-out with the man he has sat beside for 11 years with Tenerife, Valencia and now Liverpool.
The Premier League leaders are in discussion over the termination of Ayestaran's contract and an announcement is expected this week.
Benitez said: "It was a great week, we scored four and six in front of our own fans and that makes me happy. But on a personal basis I am disappointed with what's happened. He [Ayestaran] is talking to the club, but the situation will not change."
Sources at Anfield do not expect Benitez to appoint a successor. The situation came to a head during training last week at Melwood, and Ayestaran was immediately sent on leave and was not at Anfield on Saturday for the 6-0 win over Derby.
Now it is believed Benitez will take a more hands-on role in training with first-team coach Alex Miller assuming greater responsibility. Ayestaran's fitness work will be absorbed by the two fitness coaches Paco de Miguel and Antonio Gomez. Liverpool have also imported a new goalkeeping coach in Xavi Valero, who had a spell as Wrexham's keeper.
Ayestaran was in sole control of team administration and fitness training, but during the summer Benitez changed some of the training regimes in an attempt to avoid another sluggish start to the season.
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